pre-miRNA Information
pre-miRNA hsa-mir-4291   
Genomic Coordinates chr9: 93819357 - 93819421
Description Homo sapiens miR-4291 stem-loop
Comment None
RNA Secondary Structure

Mature miRNA Information
Mature miRNA hsa-miR-4291
Sequence 11| UUCAGCAGGAACAGCU |26
Evidence Experimental
Experiments SOLiD
SNPs in miRNA
Mutant ID Mutant Position Mutant Source
rs1265713434 9 dbSNP
rs1433969451 12 dbSNP
rs1372852268 16 dbSNP
Putative Targets

miRNA Expression profile
Human miRNA Tissue Atlas
Circulating MicroRNA Expression Profiling
Gene Information
Gene Symbol HEYL   
Synonyms HESR3, HEY3, HRT3, bHLHb33
Description hes related family bHLH transcription factor with YRPW motif-like
Transcript NM_014571   
Expression
Putative miRNA Targets on HEYL
3'UTR of HEYL
(miRNA target sites are highlighted)
>HEYL|NM_014571|3'UTR
   1 GCTGCCCCTTCACCACCCCGCCCCAAGGAATAAGGAAGGTTCTTTTACCAGGAGCCCAAAAAAGGGCACTGCCTTTTCTG
  81 CTTTGCTTCATGGACTGGCTCATATGTGAAGGCACGTTCTCCAGCCATCAGAGGCCCCCTCCTCCTCCAACCCATCTCTC
 161 CTTCTCACTGTTATCCCAGCTTATCCACCCAGCTCTCCTGGAGCTGTTCTGGTCTCAGAGGCTTGGTTCCATTTCTCACC
 241 TGAACAGATGAGTCCTGGGAGAGACCCTCAGAGATCCGCCCAGACCCCTCTCCTGCCCTCTGCACACCAGCAGCAGGCAT
 321 GAACCTTGGGTCTGGGAAAAAGCTTTAACCTGCAGGGCACCAGGACCCAAGGCAGGCTGTTCCTTGGGGCGGTCAGACCC
 401 CAGTCAGGAGCAATGACTGACTGGCTGCAGCCTTCCCACGCCAAGAGGCTGGAACATAGTGTCTGCCTCGCTTCCTGGAG
 481 ATAGTAACTGAGCAGGGGCTACAAAGAGGTCTCCTGGGAACCCTGTCTGCCCCTTCCCACCTGTCCTTGGGCCACACCAT
 561 CACACTGAACCACAGGACAGACCCTTTCTCCACCACAGCCAAGGCCTGGAGACTGGGGGCCCAGCAGAGCCTGCTCCCAC
 641 CCTCCTCCCAGCAGCAGACACCCACCCTCTCACTGACTAACAGGTCCCTGCACACAGCTGGCCTGGTAAACCCAGCTGGG
 721 AGGTTTCTAGGCAGCAGCAAAACTCTGTGACAGGGTGTCCTCACACCAGGCCTTGGACAGCTCTCCCAGACAGGAGCCAG
 801 GGTTGAGCAATGGAGAGCCCAGCCCCCACGTCTTACAGTCGCCATCCTCCAGGCGTGTGGTCCCTCCCCATTGGGTGCAC
 881 AGTGCAGAGGGGCCGTGGCCCCATGTGATGGTGCGCAGAGAGGAACCTCTTGGGATTCAGCACCAGACGTCTGTGCTGCC
 961 TGGTTTGCATCCGGCTCACAGAGCCCAGACTGCTGGAACAGCCAAGGACTGTCAGGCTGGACAAAAATAACTGCAAGGAG
1041 GGGCAAGAGAAAGGATGATTCGAGGCACCTTGGCCCTTCAAGGTCATGCAGTGGGTCGAGCGCCTGAGATCCTGTTCACC
1121 AGGACTCCACAGAGCTGGCTCTGCTCAGAAGCCATTTCATTCCCCGGCTCCACCCTAGGCCACTTTTTCTAACAGAGGAA
1201 ACAAATGGTCCAGCAGTCGTTCCCAGCAGAACAGCGGAGCCTGGACTGACACCCAGTGGGACCAGTGTTGCCACACCAGT
1281 TGATAAAATGCAGAAACCCTTCTGTACTCGTTGGTAAATATCTACTCCCCCAAGTGACTCCAGGTGCCCCCCACCGCCTG
1361 GCACTTCCCCCAGGACTCCTACGATCTGGTTACTGCCTGGCCGATCCAAGGCTGTGGAGTCCCAGAGCCAGCAGTTCACT
1441 GGTGCTCATTCCACACTGGTTAGATACTTCAGTTGTCACCCCTGGGAAGATTCTCCCACCTCCTCCCTTTGATGGAACCA
1521 CCCTCCCCAGAGGCTGCATTGAGGAGACTCCACAGACTGAAAAGTGAGTTTGCAGAAACCTTGGGGAAAAGGGCCCTTTC
1601 AAAGAAGTGGATAAGAGGGAGGAGATCATTGAGTGACCCAGAAAGCTCTTTTGAAAAGACAGACTCCTCAAGGAGAGATA
1681 AAGAGGAAAGCACCTCTTTCATTTTTTAGTGTGAGCTAATTCCATCAGACTGCTGTCCTCCTGGACCCATCTGAGATGTG
1761 CAGTAGCAAGGAGAGGGGGGATCATTTTAGAGAGTGGGTCATTGGCAGGGAGTGCTCCGGAGGGAGGCAGAGGGGAGACT
1841 GTGGTAGAAGGAAGACAGAACTCACACATGCTCCCAGGATTGGGGACAGGGACAGAGGAGGTAACAGAAGGCAAAGGCCA
1921 GTTTCCCCGTTATCATGAAGGGGCCCACTCAGGACAGGAACAAGGACAACTCCTCCTCCTCCTCCTCCTCTCCTGCTGCT
2001 CCTGGGATACCAGGTCAGTGATGTAGTCTTGCAGTTTGGCAACTTCCTAGCCTGAGAATCCCTAGTGGGGCTGTGGGAAA
2081 CACATTTCCACGTTGCAAGCATGCAACTCCAAAGAATCTGTGATGCCACTGAAATGAGATGGGAATGATCCAGCTCTTTC
2161 AGCATCTTGGTTGAACTTGCTTTCATTGTCCCTGGGATATTGTGGAAGGAAAGGTGACTGTGTGATCTGATTCTGTGGTC
2241 AAGGACTTGCATCTTGTGTTTCTATCCCCAAGCCTTCCTGGTGTCTCCAACTCCTACCCCATTGCATGGGTTGTTGCGGA
2321 CATCCAATAAAGATTTTTTTAGTGCTTCTGGAAACTTCCAGTAGATTCTACTTCTAAACTATCTCTGGAGTCCATCCACT
2401 TCTGTCTGCACCCACAGCCATCCTGGCCAGGCCACATCACCTCCCCCAGATCACTGCCCTGGCCTCAGAAAGGTCTTCCC
2481 TCTTGCTTTGTCAATCAGTTCTCAGTAGCAGCAGAGAGAAATTGAAAGCTGCAGGTCATATCGTATCATCTTTAGTTTGA
2561 AAACCTCACTCTCTTACCCTATTGTTCTAAAGGTCTTCTTTTGGTCCCAACCTCATTTCCAGCCTCATTTCTTGCCAGTC
2641 CCAGACTTGCTCCCTGAGCTTCTGCCACCTGCCCTTCCTTCATTTCCTCGACATTCCAGCCTTGTTCCCACCTCCAGCAC
2721 TTTGCATATGCTGTTCCCTTTGCCAAGAATGCTCTTCCCCTACCCTGTGCATGGCTGAGTTCTGCAGACCCTCAGGCCTT
2801 GGCTTCAACGTTGCCTCGTCCAAGAGGCCTTCCTCGACTACTTTACTTGTGGAGTTCCTCTATCACAAGGCCTCTGTTCT
2881 TTCCCTTCATGGAGAATTTGCCACTGCATATCCATTTGTGTAATTTACTTGTTGGTTGACTGTGCCTCCCACTCGAGTGT
2961 AAGCTCATGAGGCCAGGTGCCATGCCTGGTTCAGTCTCCACTCTGTACCCAGCATTGAGCACAGGGCCTGGTCCATAGTT
3041 GGCGTTCAATAAATACTTGTTGAAGAAGTGAACTGAAAAAAAAAAAAAAAAAA
Target sites Provided by authors   Predicted by miRanda    DRVs    SNPs    DRVs & SNPs
miRNA-target interactions
(Predicted by miRanda)
ID Duplex structure Position Score MFE
1
miRNA  3' ucgacaAGGACGACuu 5'
                ||||||||  
Target 5' ctcctcTCCTGCTGct 3'
1985 - 2000 130.00 -11.70
2
miRNA  3' ucgacaagGACGACUu 5'
                  ||||||: 
Target 5' agcccagaCTGCTGGa 3'
982 - 997 124.00 -12.00
3
miRNA  3' ucgacaagGACGACuu 5'
                  ||||||  
Target 5' ccatcagaCTGCTGtc 3'
1722 - 1737 120.00 -8.70
DRVs in gene 3'UTRs
Mutant ID Mutant Position Mutant Source
COSN30167361 28 COSMIC
COSN31519122 42 COSMIC
COSN30140067 140 COSMIC
COSN30531403 167 COSMIC
COSN28785822 199 COSMIC
COSN16365422 246 COSMIC
COSN19161846 439 COSMIC
COSN8469177 459 COSMIC
COSN22671896 778 COSMIC
COSN15086001 860 COSMIC
COSN19552206 1227 COSMIC
COSN7219363 1356 COSMIC
COSN7219362 1524 COSMIC
COSN2531136 1970 COSMIC
COSN25428678 2339 COSMIC
rs116401702 2754 GWAS
SNPs in gene 3'UTRs
Mutant ID Mutant Position Mutant Source
rs1292797893 2 dbSNP
rs1433961695 5 dbSNP
rs772575144 7 dbSNP
rs1321668570 8 dbSNP
rs764222546 11 dbSNP
rs1443096054 12 dbSNP
rs183890301 13 dbSNP
rs191371215 16 dbSNP
rs759910471 19 dbSNP
rs1370321375 20 dbSNP
rs150591457 24 dbSNP
rs996221201 28 dbSNP
rs904529912 39 dbSNP
rs1205940809 41 dbSNP
rs1235836667 42 dbSNP
rs1438343359 44 dbSNP
rs1042937923 46 dbSNP
rs1449722106 51 dbSNP
rs1006227707 56 dbSNP
rs945970122 58 dbSNP
rs1426248902 63 dbSNP
rs550856748 66 dbSNP
rs531270507 67 dbSNP
rs1413268739 69 dbSNP
rs1410749906 71 dbSNP
rs1330926974 76 dbSNP
rs913033407 78 dbSNP
rs1398226470 85 dbSNP
rs527386529 86 dbSNP
rs1048461291 89 dbSNP
rs2695320 90 dbSNP
rs750013021 95 dbSNP
rs1215996385 101 dbSNP
rs902032969 104 dbSNP
rs1198537339 106 dbSNP
rs1244355445 107 dbSNP
rs1444648503 111 dbSNP
rs1185781164 113 dbSNP
rs938415414 115 dbSNP
rs927083349 116 dbSNP
rs1173152270 120 dbSNP
rs979729983 122 dbSNP
rs528479111 135 dbSNP
rs765261851 137 dbSNP
rs1207467707 151 dbSNP
rs559450152 155 dbSNP
rs930947034 161 dbSNP
rs1177058068 162 dbSNP
rs1040508977 163 dbSNP
rs1394226337 173 dbSNP
rs115608339 174 dbSNP
rs1432431726 175 dbSNP
rs143702140 177 dbSNP
rs1316732799 179 dbSNP
rs1223532078 180 dbSNP
rs1264461957 183 dbSNP
rs1347704755 184 dbSNP
rs188142662 186 dbSNP
rs1255875835 189 dbSNP
rs924946962 190 dbSNP
rs1015081753 206 dbSNP
rs1337340002 225 dbSNP
rs988071220 226 dbSNP
rs954775826 229 dbSNP
rs1226933869 233 dbSNP
rs548817491 241 dbSNP
rs183266275 248 dbSNP
rs1379480700 254 dbSNP
rs1464441106 262 dbSNP
rs574422527 267 dbSNP
rs1171978657 277 dbSNP
rs554431129 278 dbSNP
rs1402299761 282 dbSNP
rs1296127142 288 dbSNP
rs1025261565 289 dbSNP
rs540895508 292 dbSNP
rs1383920660 299 dbSNP
rs996263985 300 dbSNP
rs1284589708 303 dbSNP
rs991989362 305 dbSNP
rs1231667705 309 dbSNP
rs1255989361 310 dbSNP
rs959154514 313 dbSNP
rs1219179184 314 dbSNP
rs41264501 324 dbSNP
rs1284335971 325 dbSNP
rs558220549 349 dbSNP
rs1006281696 354 dbSNP
rs1431552639 357 dbSNP
rs1325073628 363 dbSNP
rs201450931 368 dbSNP
rs2307917 368 dbSNP
rs552280913 390 dbSNP
rs1026297603 391 dbSNP
rs1165840994 399 dbSNP
rs994184708 401 dbSNP
rs867546198 408 dbSNP
rs538737122 413 dbSNP
rs569642442 422 dbSNP
rs753664881 428 dbSNP
rs1373212313 439 dbSNP
rs376779376 440 dbSNP
rs1040561492 443 dbSNP
rs1385786618 449 dbSNP
rs1307106237 457 dbSNP
rs1336805096 459 dbSNP
rs1252943078 467 dbSNP
rs763815500 470 dbSNP
rs1056962089 472 dbSNP
rs557578449 489 dbSNP
rs1223467293 490 dbSNP
rs1214816834 492 dbSNP
rs1267526643 493 dbSNP
rs943545683 495 dbSNP
rs889251636 499 dbSNP
rs1251375062 503 dbSNP
rs938552211 517 dbSNP
rs137911802 519 dbSNP
rs879401511 519 dbSNP
rs190993934 523 dbSNP
rs977777418 530 dbSNP
rs945324513 538 dbSNP
rs1366287416 541 dbSNP
rs1044086727 546 dbSNP
rs1272081222 559 dbSNP
rs563024194 565 dbSNP
rs1329961019 573 dbSNP
rs931082575 575 dbSNP
rs1231180254 584 dbSNP
rs375847567 588 dbSNP
rs919574677 619 dbSNP
rs1244361668 620 dbSNP
rs1302620493 637 dbSNP
rs1459173366 644 dbSNP
rs755620716 645 dbSNP
rs959206466 648 dbSNP
rs1461720565 650 dbSNP
rs1033490156 670 dbSNP
rs1372987004 675 dbSNP
rs1326599489 676 dbSNP
rs1422834113 679 dbSNP
rs1409084754 686 dbSNP
rs1425112266 690 dbSNP
rs972841278 692 dbSNP
rs984898457 694 dbSNP
rs1169152391 698 dbSNP
rs940937276 708 dbSNP
rs1400136390 714 dbSNP
rs760908899 717 dbSNP
rs952151491 727 dbSNP
rs1377497963 728 dbSNP
rs908161200 728 dbSNP
rs775579059 731 dbSNP
rs1026769472 737 dbSNP
rs954817971 738 dbSNP
rs1320070790 749 dbSNP
rs771391854 750 dbSNP
rs1257506668 754 dbSNP
rs993512141 760 dbSNP
rs1212183551 791 dbSNP
rs1029181979 794 dbSNP
rs568805127 795 dbSNP
rs1019166182 798 dbSNP
rs1455958276 799 dbSNP
rs767575178 811 dbSNP
rs969235040 812 dbSNP
rs1476644093 813 dbSNP
rs1021608771 822 dbSNP
rs1007739955 825 dbSNP
rs147241011 827 dbSNP
rs1178918745 828 dbSNP
rs1438695121 829 dbSNP
rs1275411250 830 dbSNP
rs1198905984 836 dbSNP
rs759594991 840 dbSNP
rs774475980 841 dbSNP
rs528765186 846 dbSNP
rs905682328 847 dbSNP
rs1044222373 855 dbSNP
rs1441234109 861 dbSNP
rs188708237 862 dbSNP
rs552246769 865 dbSNP
rs559685317 866 dbSNP
rs1487075380 870 dbSNP
rs945069388 871 dbSNP
rs771393922 872 dbSNP
rs1264189012 880 dbSNP
rs1478763168 888 dbSNP
rs898264818 894 dbSNP
rs1423607081 895 dbSNP
rs749839999 895 dbSNP
rs1159585025 897 dbSNP
rs937893430 899 dbSNP
rs1412518998 904 dbSNP
rs532688278 913 dbSNP
rs1418076615 914 dbSNP
rs1374386250 915 dbSNP
rs1189537248 920 dbSNP
rs376278740 925 dbSNP
rs1438773539 927 dbSNP
rs1273167175 930 dbSNP
rs1366742897 938 dbSNP
rs939717328 948 dbSNP
rs984634727 949 dbSNP
rs549601918 967 dbSNP
rs1406444363 969 dbSNP
rs912231052 970 dbSNP
rs143934134 972 dbSNP
rs1290214336 973 dbSNP
rs1470735179 979 dbSNP
rs1234929598 983 dbSNP
rs1218652497 984 dbSNP
rs560722152 988 dbSNP
rs148112227 992 dbSNP
rs1488414466 997 dbSNP
rs922079524 1000 dbSNP
rs1382527527 1003 dbSNP
rs1438305382 1006 dbSNP
rs974930238 1023 dbSNP
rs1485464777 1029 dbSNP
rs1258406690 1030 dbSNP
rs1414730681 1035 dbSNP
rs968828606 1040 dbSNP
rs1351534720 1058 dbSNP
rs1321628323 1059 dbSNP
rs11206478 1060 dbSNP
rs564937424 1061 dbSNP
rs953620522 1062 dbSNP
rs956196233 1068 dbSNP
rs1281064225 1073 dbSNP
rs1035673392 1074 dbSNP
rs1002716050 1097 dbSNP
rs1318053157 1098 dbSNP
rs1313048593 1105 dbSNP
rs1010866304 1109 dbSNP
rs1240639765 1121 dbSNP
rs969977461 1129 dbSNP
rs1456834277 1135 dbSNP
rs1181454968 1138 dbSNP
rs1238445425 1139 dbSNP
rs544674309 1151 dbSNP
rs1473871326 1152 dbSNP
rs576115030 1154 dbSNP
rs1466261616 1159 dbSNP
rs1042152735 1160 dbSNP
rs560538573 1163 dbSNP
rs182773043 1164 dbSNP
rs536317655 1165 dbSNP
rs879044838 1166 dbSNP
rs926526778 1172 dbSNP
rs1003982545 1178 dbSNP
rs890757263 1181 dbSNP
rs930503451 1189 dbSNP
rs1410703787 1197 dbSNP
rs919058643 1202 dbSNP
rs1163229237 1206 dbSNP
rs1219874138 1208 dbSNP
rs1255613631 1209 dbSNP
rs1050775058 1218 dbSNP
rs575104511 1219 dbSNP
rs1190276808 1230 dbSNP
rs922218853 1235 dbSNP
rs1484954539 1236 dbSNP
rs1241568270 1239 dbSNP
rs986091098 1240 dbSNP
rs767088418 1246 dbSNP
rs1039212580 1247 dbSNP
rs117679560 1250 dbSNP
rs947506745 1253 dbSNP
rs914748180 1256 dbSNP
rs540325312 1280 dbSNP
rs12049221 1281 dbSNP
rs1020853505 1297 dbSNP
rs928627060 1302 dbSNP
rs981432631 1310 dbSNP
rs1009348183 1319 dbSNP
rs1234713873 1320 dbSNP
rs1401496643 1322 dbSNP
rs566231366 1329 dbSNP
rs778617301 1333 dbSNP
rs1275705274 1337 dbSNP
rs143683714 1341 dbSNP
rs1227332994 1344 dbSNP
rs1276020293 1345 dbSNP
rs1034723971 1347 dbSNP
rs1269072580 1352 dbSNP
rs552571643 1355 dbSNP
rs904827548 1356 dbSNP
rs538642690 1359 dbSNP
rs1488709973 1365 dbSNP
rs1219131227 1366 dbSNP
rs148982776 1367 dbSNP
rs41264499 1374 dbSNP
rs1199479355 1378 dbSNP
rs930535569 1381 dbSNP
rs897684277 1382 dbSNP
rs1172741471 1383 dbSNP
rs1360957481 1388 dbSNP
rs1421119791 1389 dbSNP
rs1004354315 1402 dbSNP
rs530089320 1403 dbSNP
rs911979078 1416 dbSNP
rs138871678 1426 dbSNP
rs777557147 1427 dbSNP
rs1333290449 1433 dbSNP
rs931996909 1435 dbSNP
rs750193277 1439 dbSNP
rs1303098072 1446 dbSNP
rs1418984078 1448 dbSNP
rs1050745926 1450 dbSNP
rs925922531 1451 dbSNP
rs996547084 1455 dbSNP
rs899508326 1457 dbSNP
rs1039263673 1463 dbSNP
rs1187193111 1465 dbSNP
rs1485675305 1468 dbSNP
rs1210444735 1472 dbSNP
rs1269599039 1473 dbSNP
rs1449748994 1481 dbSNP
rs1476401644 1487 dbSNP
rs1192986625 1490 dbSNP
rs1380650655 1505 dbSNP
rs1156627597 1525 dbSNP
rs979060619 1527 dbSNP
rs1458119957 1543 dbSNP
rs547181648 1547 dbSNP
rs1388006526 1579 dbSNP
rs1439300401 1581 dbSNP
rs968022278 1583 dbSNP
rs1020485859 1592 dbSNP
rs1467899961 1593 dbSNP
rs766204608 1594 dbSNP
rs1273556921 1599 dbSNP
rs1223400152 1604 dbSNP
rs1245829663 1605 dbSNP
rs527632820 1611 dbSNP
rs1354383400 1622 dbSNP
rs1271777733 1628 dbSNP
rs1238652191 1629 dbSNP
rs755874286 1629 dbSNP
rs1176497844 1632 dbSNP
rs565009299 1637 dbSNP
rs1424007214 1643 dbSNP
rs1301931935 1657 dbSNP
rs1366195442 1659 dbSNP
rs1460520117 1667 dbSNP
rs545021876 1668 dbSNP
rs575964222 1669 dbSNP
rs1401577196 1670 dbSNP
rs562437127 1679 dbSNP
rs1296765902 1681 dbSNP
rs1373122273 1685 dbSNP
rs981401694 1691 dbSNP
rs1372251142 1693 dbSNP
rs1396797590 1698 dbSNP
rs1315189311 1700 dbSNP
rs372407957 1700 dbSNP
rs1328346859 1709 dbSNP
rs1232677701 1724 dbSNP
rs1258826772 1731 dbSNP
rs752592131 1738 dbSNP
rs948691148 1741 dbSNP
rs1196725621 1743 dbSNP
rs904880900 1744 dbSNP
rs915917463 1747 dbSNP
rs1027718465 1749 dbSNP
rs974366388 1756 dbSNP
rs41264497 1757 dbSNP
rs1015823223 1769 dbSNP
rs535251181 1770 dbSNP
rs575868941 1771 dbSNP
rs1029383131 1775 dbSNP
rs1393315606 1776 dbSNP
rs555240800 1779 dbSNP
rs944549576 1785 dbSNP
rs1466817717 1798 dbSNP
rs1304962686 1802 dbSNP
rs534886445 1803 dbSNP
rs1219474137 1811 dbSNP
rs1191722501 1814 dbSNP
rs899475310 1818 dbSNP
rs190986521 1819 dbSNP
rs1192719510 1828 dbSNP
rs1479507103 1831 dbSNP
rs1278897414 1834 dbSNP
rs559005718 1835 dbSNP
rs1012229269 1840 dbSNP
rs543421470 1848 dbSNP
rs932049436 1852 dbSNP
rs1053694126 1855 dbSNP
rs1312716877 1869 dbSNP
rs867901099 1873 dbSNP
rs539294029 1876 dbSNP
rs934866341 1880 dbSNP
rs1277550749 1881 dbSNP
rs907390903 1890 dbSNP
rs1234290186 1896 dbSNP
rs1485112469 1901 dbSNP
rs1216954038 1902 dbSNP
rs1264672855 1903 dbSNP
rs1185277112 1909 dbSNP
rs1476595039 1909 dbSNP
rs772082951 1928 dbSNP
rs1045755551 1929 dbSNP
rs1178620114 1933 dbSNP
rs948826589 1935 dbSNP
rs1349245719 1936 dbSNP
rs913478370 1937 dbSNP
rs539771599 1944 dbSNP
rs1299102356 1975 dbSNP
rs1390053466 1978 dbSNP
rs960209854 1979 dbSNP
rs927423697 1980 dbSNP
rs1288368740 1984 dbSNP
rs974123807 1987 dbSNP
rs1220894884 1991 dbSNP
rs1344264175 1991 dbSNP
rs879122375 1991 dbSNP
rs980542506 1991 dbSNP
rs186178580 2005 dbSNP
rs1373771551 2006 dbSNP
rs1173093734 2014 dbSNP
rs569718016 2018 dbSNP
rs1395635774 2022 dbSNP
rs1178091376 2024 dbSNP
rs941270996 2036 dbSNP
rs1417120217 2039 dbSNP
rs1027360978 2059 dbSNP
rs1489078205 2061 dbSNP
rs1177482037 2063 dbSNP
rs1483828059 2067 dbSNP
rs1454153191 2068 dbSNP
rs745811962 2072 dbSNP
rs549724585 2077 dbSNP
rs1399208589 2091 dbSNP
rs1485687278 2092 dbSNP
rs536388733 2096 dbSNP
rs1237480051 2098 dbSNP
rs778818220 2104 dbSNP
rs375034027 2106 dbSNP
rs1311221883 2117 dbSNP
rs1316747699 2117 dbSNP
rs11554176 2127 dbSNP
rs41264495 2136 dbSNP
rs975131301 2145 dbSNP
rs1289928725 2146 dbSNP
rs890179738 2152 dbSNP
rs1216081596 2163 dbSNP
rs763387335 2164 dbSNP
rs963817437 2166 dbSNP
rs1022361138 2172 dbSNP
rs547216509 2173 dbSNP
rs41264493 2180 dbSNP
rs1415509679 2183 dbSNP
rs41264491 2187 dbSNP
rs1043268038 2189 dbSNP
rs1176884544 2190 dbSNP
rs999067758 2191 dbSNP
rs907359987 2192 dbSNP
rs946282069 2200 dbSNP
rs913511442 2201 dbSNP
rs375071116 2204 dbSNP
rs1162895706 2216 dbSNP
rs1363078174 2221 dbSNP
rs6587 2234 dbSNP
rs1012934569 2235 dbSNP
rs927330382 2242 dbSNP
rs548338263 2253 dbSNP
rs1284484387 2256 dbSNP
rs531584592 2262 dbSNP
rs1038767595 2280 dbSNP
rs1230850347 2282 dbSNP
rs920417556 2283 dbSNP
rs973291099 2285 dbSNP
rs941336820 2286 dbSNP
rs1199878581 2291 dbSNP
rs1275516881 2301 dbSNP
rs962252286 2302 dbSNP
rs1257635021 2305 dbSNP
rs1238211877 2306 dbSNP
rs1014812937 2308 dbSNP
rs1203955672 2314 dbSNP
rs78706896 2317 dbSNP
rs542633090 2318 dbSNP
rs1046990980 2319 dbSNP
rs933907944 2321 dbSNP
rs922417006 2327 dbSNP
rs1409095346 2330 dbSNP
rs1334569355 2331 dbSNP
rs996325890 2334 dbSNP
rs1375001585 2340 dbSNP
rs1412564938 2341 dbSNP
rs1292109982 2348 dbSNP
rs975267919 2351 dbSNP
rs963787313 2371 dbSNP
rs1272492046 2375 dbSNP
rs915031217 2376 dbSNP
rs1214285365 2380 dbSNP
rs1342397426 2381 dbSNP
rs1279806898 2383 dbSNP
rs989644741 2384 dbSNP
rs956361668 2397 dbSNP
rs1031106054 2398 dbSNP
rs180995630 2408 dbSNP
rs529043487 2432 dbSNP
rs1043602900 2435 dbSNP
rs1256889519 2440 dbSNP
rs1488691671 2446 dbSNP
rs560276047 2447 dbSNP
rs971642292 2449 dbSNP
rs1024423132 2465 dbSNP
rs1013070542 2466 dbSNP
rs540291443 2471 dbSNP
rs1172897754 2472 dbSNP
rs771830719 2478 dbSNP
rs1432445115 2485 dbSNP
rs1311964595 2486 dbSNP
rs1321088962 2494 dbSNP
rs1458650730 2501 dbSNP
rs1391979512 2504 dbSNP
rs140314362 2509 dbSNP
rs752337633 2511 dbSNP
rs1472914688 2521 dbSNP
rs1315619305 2522 dbSNP
rs1369764845 2524 dbSNP
rs1190404722 2531 dbSNP
rs559065410 2537 dbSNP
rs1057401492 2541 dbSNP
rs938627605 2542 dbSNP
rs545387557 2543 dbSNP
rs1350580905 2544 dbSNP
rs531931512 2555 dbSNP
rs1279810202 2564 dbSNP
rs1044451515 2569 dbSNP
rs1206929600 2571 dbSNP
rs1259771858 2575 dbSNP
rs576319266 2579 dbSNP
rs745539827 2581 dbSNP
rs887086897 2582 dbSNP
rs12030495 2584 dbSNP
rs770567619 2590 dbSNP
rs940548085 2599 dbSNP
rs796152277 2604 dbSNP
rs907763918 2610 dbSNP
rs1383665703 2611 dbSNP
rs1382148621 2614 dbSNP
rs2695321 2616 dbSNP
rs1358740876 2620 dbSNP
rs200913639 2635 dbSNP
rs552545622 2637 dbSNP
rs1327063868 2650 dbSNP
rs1298297113 2652 dbSNP
rs1377861350 2663 dbSNP
rs151170453 2689 dbSNP
rs1039550500 2690 dbSNP
rs942516124 2696 dbSNP
rs1368588886 2702 dbSNP
rs534271665 2703 dbSNP
rs1324970773 2707 dbSNP
rs756140334 2710 dbSNP
rs1405673071 2711 dbSNP
rs1350132174 2712 dbSNP
rs1210347918 2718 dbSNP
rs142093126 2720 dbSNP
rs868518119 2737 dbSNP
rs1467090601 2739 dbSNP
rs1192735305 2743 dbSNP
rs989223898 2745 dbSNP
rs1420468992 2750 dbSNP
rs116401702 2754 dbSNP
rs1362690710 2759 dbSNP
rs1461079924 2760 dbSNP
rs114297514 2771 dbSNP
rs1397275105 2774 dbSNP
rs1021390597 2779 dbSNP
rs1010789277 2783 dbSNP
rs891974296 2790 dbSNP
rs1453108202 2808 dbSNP
rs12022423 2809 dbSNP
rs765821510 2814 dbSNP
rs971612562 2817 dbSNP
rs755140248 2818 dbSNP
rs372439881 2823 dbSNP
rs1013124036 2831 dbSNP
rs1481525321 2833 dbSNP
rs958877316 2835 dbSNP
rs1017533430 2836 dbSNP
rs1424060255 2840 dbSNP
rs1044774584 2842 dbSNP
rs1415086988 2846 dbSNP
rs549035554 2854 dbSNP
rs528912772 2860 dbSNP
rs560162992 2862 dbSNP
rs1195416825 2865 dbSNP
rs1355467994 2882 dbSNP
rs898546120 2883 dbSNP
rs1276849469 2885 dbSNP
rs1037126839 2889 dbSNP
rs539955756 2894 dbSNP
rs907711590 2906 dbSNP
rs533007649 2914 dbSNP
rs1314810640 2919 dbSNP
rs762933616 2926 dbSNP
rs1353711787 2931 dbSNP
rs987299045 2935 dbSNP
rs1436968404 2941 dbSNP
rs1025648146 2949 dbSNP
rs998222536 2950 dbSNP
rs1309186880 2954 dbSNP
rs901182943 2955 dbSNP
rs1390447742 2958 dbSNP
rs1483360381 2964 dbSNP
rs1212627068 2966 dbSNP
rs565382100 2967 dbSNP
rs1254772012 2984 dbSNP
rs1370704150 2996 dbSNP
rs974789585 2999 dbSNP
rs1039603740 3008 dbSNP
rs968424492 3009 dbSNP
rs1430844435 3015 dbSNP
rs1168721517 3019 dbSNP
rs189693750 3024 dbSNP
rs765632241 3030 dbSNP
rs893613699 3032 dbSNP
rs1054059509 3035 dbSNP
rs1315073773 3036 dbSNP
rs1373880430 3039 dbSNP
rs988578140 3043 dbSNP
rs955809876 3044 dbSNP
rs1380507541 3045 dbSNP
rs1294547050 3058 dbSNP
rs1348408253 3059 dbSNP
rs1236025126 3065 dbSNP
rs935090693 3068 dbSNP
Experimental Support 1 for Functional miRNA-Target Interaction
miRNA:Target ----
Validation Method
Conditions LCL-BAC-D2
Location of target site 3'UTR
Tools used in this research TargetScan , miRTarCLIP , Piranha
Original Description (Extracted from the article) ... PAR-CLIP data was present in GSM1133252. RNA binding protein: AGO2. Condition:Untreated ...

- Majoros WH; Lekprasert P; Mukherjee N; et al., 2013, Nature methods.

miRNA-target interactions (Provided by authors)
ID Duplex structure Position
1
miRNA  3' ucgacaagGACGACUu 5'
                  ||||||| 
Target 5' ---uugggCUGCUGAg 3'
1 - 13
Article - Majoros WH; Lekprasert P; Mukherjee N; et al.
- Nature methods, 2013
High-throughput sequencing has opened numerous possibilities for the identification of regulatory RNA-binding events. Cross-linking and immunoprecipitation of Argonaute proteins can pinpoint a microRNA (miRNA) target site within tens of bases but leaves the identity of the miRNA unresolved. A flexible computational framework, microMUMMIE, integrates sequence with cross-linking features and reliably identifies the miRNA family involved in each binding event. It considerably outperforms sequence-only approaches and quantifies the prevalence of noncanonical binding modes.
LinkOut: [PMID: 23708386]
CLIP-seq Support 1 for dataset GSM1133252
Method / RBP PAR-CLIP / AGO2
Cell line / Condition LCL-BAC-D2 / Untreated
Location of target site ENST00000372852.3 | 3UTR | UUGGGCUGCUGAGAG
Tools used in this analysis TargetScan, miRTarCLIP, and Piranha
Article / Accession Series PMID: 23708386 / GSE46611
CLIP-seq Viewer Link
MiRNA-Target Expression Profile
Dataset Pearson Correlation P-value for Pearson Correlation Spearman Correlation P-value for Spearman Correlation Samples Chart
MiRNA-Target Expression Profile (TCGA)
Tumor Pearson Correlation P-value for Pearson Correlation Spearman Correlation P-value for Spearman Correlation Samples Chart
82 hsa-miR-4291 Target Genes:
Functional analysis:
ID Target Description Validation methods
Strong evidence Less strong evidence
MIRT102445 CALU calumenin 2 4
MIRT108677 ZBTB33 zinc finger and BTB domain containing 33 2 4
MIRT125969 SHOC2 SHOC2, leucine rich repeat scaffold protein 2 6
MIRT179018 PAFAH1B2 platelet activating factor acetylhydrolase 1b catalytic subunit 2 2 4
MIRT379033 CDK6 cyclin dependent kinase 6 2 6
MIRT442473 CPEB4 cytoplasmic polyadenylation element binding protein 4 2 2
MIRT442910 PCBD2 pterin-4 alpha-carbinolamine dehydratase 2 2 2
MIRT442979 ZNF736 zinc finger protein 736 2 2
MIRT445663 TNFSF15 TNF superfamily member 15 2 2
MIRT446237 FZD6 frizzled class receptor 6 2 2
MIRT448860 FAM49B family with sequence similarity 49 member B 2 2
MIRT455559 TRAF1 TNF receptor associated factor 1 2 2
MIRT459704 ZNF641 zinc finger protein 641 2 2
MIRT460608 FEM1A fem-1 homolog A 2 2
MIRT462251 LAMA4 laminin subunit alpha 4 2 2
MIRT462469 FIZ1 FLT3 interacting zinc finger 1 2 2
MIRT466656 TAF1D TATA-box binding protein associated factor, RNA polymerase I subunit D 2 6
MIRT466841 STX6 syntaxin 6 2 2
MIRT471403 PDP2 pyruvate dehyrogenase phosphatase catalytic subunit 2 2 2
MIRT471517 PCGF3 polycomb group ring finger 3 2 2
MIRT471681 PABPN1 poly(A) binding protein nuclear 1 2 2
MIRT472858 MTHFD2 methylenetetrahydrofolate dehydrogenase (NADP+ dependent) 2, methenyltetrahydrofolate cyclohydrolase 2 2
MIRT474813 KIAA0226 RUN and cysteine rich domain containing beclin 1 interacting protein 2 2
MIRT474931 KCTD15 potassium channel tetramerization domain containing 15 2 2
MIRT475814 HDGF heparin binding growth factor 2 2
MIRT480434 C17orf49 chromosome 17 open reading frame 49 2 2
MIRT480903 BCL2L2-PABPN1 BCL2L2-PABPN1 readthrough 2 2
MIRT481478 ARL8B ADP ribosylation factor like GTPase 8B 2 2
MIRT484982 UBE2V1 ubiquitin conjugating enzyme E2 V1 2 8
MIRT485019 TMEM189-UBE2V1 TMEM189-UBE2V1 readthrough 2 8
MIRT485036 TMEM189 transmembrane protein 189 2 8
MIRT495074 HEYL hes related family bHLH transcription factor with YRPW motif-like 2 2
MIRT496004 CD180 CD180 molecule 2 2
MIRT500675 TRIM37 tripartite motif containing 37 2 2
MIRT504544 ZNF417 zinc finger protein 417 2 6
MIRT506782 KLHL15 kelch like family member 15 2 6
MIRT507256 FGF2 fibroblast growth factor 2 2 6
MIRT507386 EN2 engrailed homeobox 2 2 2
MIRT512505 BTBD19 BTB domain containing 19 2 2
MIRT516903 CTSB cathepsin B 2 2
MIRT528124 PPP1R10 protein phosphatase 1 regulatory subunit 10 2 2
MIRT528874 ATF3 activating transcription factor 3 2 2
MIRT536091 MBOAT2 membrane bound O-acyltransferase domain containing 2 2 2
MIRT541079 RLIM ring finger protein, LIM domain interacting 2 2
MIRT541099 RAF1 Raf-1 proto-oncogene, serine/threonine kinase 2 2
MIRT545360 LIN7C lin-7 homolog C, crumbs cell polarity complex component 2 2
MIRT545831 ZNF367 zinc finger protein 367 2 4
MIRT547115 PHLPP2 PH domain and leucine rich repeat protein phosphatase 2 2 2
MIRT547312 NPTN neuroplastin 2 2
MIRT547959 HIGD1A HIG1 hypoxia inducible domain family member 1A 2 4
MIRT549943 RPL7L1 ribosomal protein L7 like 1 2 2
MIRT550749 ENTPD1 ectonucleoside triphosphate diphosphohydrolase 1 2 2
MIRT565145 TUBB2A tubulin beta 2A class IIa 2 2
MIRT571050 POLQ DNA polymerase theta 2 2
MIRT571361 ZNF45 zinc finger protein 45 2 2
MIRT610133 FOXI2 forkhead box I2 2 2
MIRT613145 DSE dermatan sulfate epimerase 2 2
MIRT613379 ABCC12 ATP binding cassette subfamily C member 12 2 2
MIRT615752 C6 complement C6 2 2
MIRT616460 ADRA2B adrenoceptor alpha 2B 2 2
MIRT616719 FEM1B fem-1 homolog B 2 2
MIRT618312 IPP intracisternal A particle-promoted polypeptide 2 2
MIRT631491 RASSF4 Ras association domain family member 4 2 2
MIRT643134 PLCXD2 phosphatidylinositol specific phospholipase C X domain containing 2 2 2
MIRT643482 DISC1 disrupted in schizophrenia 1 2 2
MIRT649715 TWSG1 twisted gastrulation BMP signaling modulator 1 2 2
MIRT653542 SLC38A7 solute carrier family 38 member 7 2 2
MIRT666307 SLC22A3 solute carrier family 22 member 3 2 2
MIRT692005 NAP1L4 nucleosome assembly protein 1 like 4 2 2
MIRT696838 ARL2BP ADP ribosylation factor like GTPase 2 binding protein 2 2
MIRT698204 TMEM248 transmembrane protein 248 2 2
MIRT703316 GDPD5 glycerophosphodiester phosphodiesterase domain containing 5 2 2
MIRT709376 FAM13A family with sequence similarity 13 member A 2 2
MIRT710112 MED23 mediator complex subunit 23 2 2
MIRT711975 HOMER2 homer scaffolding protein 2 2 2
MIRT712275 PPIP5K2 diphosphoinositol pentakisphosphate kinase 2 2 2
MIRT714108 TMED9 transmembrane p24 trafficking protein 9 2 2
MIRT719113 MAML1 mastermind like transcriptional coactivator 1 2 2
MIRT719727 SLC39A11 solute carrier family 39 member 11 2 2
MIRT720018 TFAP2C transcription factor AP-2 gamma 2 2
MIRT720358 ZBTB8B zinc finger and BTB domain containing 8B 2 2
MIRT720372 NUDT3 nudix hydrolase 3 2 2
miRNA-Drug Associations
miRNA Small Melocule FDA CID Detection Method Condition PMID Year Expression Pattern of miRNA
miR-4 Dexamethasone approved 5743 Microarray primary rat thymocytes 20847043 2010 up-regulated
miRNA-Drug Resistance Associations
miRNA Drug Name CID NSC FDA Effect/Pattern Detection Method Level Phenotype Condition
hsa-miR-4291 Platinum-based doublet chemotherapy resistant High Lung Adenocarcinoma tissue
hsa-miR-4291 Platinum 23939 resistant High Ovarian Cancer tissue
hsa-miR-4291 Paclitaxel 36314 NSC125973 approved resistant High Prostate Cancer cell line (DU-145, PC-3)
hsa-miR-4291 Doxorubicin 31703 NSC123127 approved resistant High Chronic Myelogenous Leukemia cell line (K562)
hsa-miR-4291 Radioactivity Iodine resistant High Papillary Thyroid Cancer tissue
hsa-miR-4291 Cisplatin 5460033 NSC119875 approved resistant High Hypopharyngeal Cancer cell line (FaDu)
hsa-miR-4291 Gefitinib 123631 NSC715055 approved resistant cell line (PC9)
hsa-miR-4291 Osimertinib 71496458 NSC779217 approved resistant cell line (PC9)
hsa-miR-4291 Cisplatin 5460033 NSC119875 approved sensitive cell line (CAL-27) (cytosolic RNA)
hsa-miR-4291 Osimertinib 71496458 NSC779217 approved sensitive cell line (H1975)
hsa-miR-4291 Platinum-based doublet chemotherapy resistant tissue (lung adenocarcinoma)
hsa-miR-4291 Cisplatin 5460033 NSC119875 approved sensitive cell line (A549)
hsa-miR-4291 Gemcitabine 60750 NSC613327 approved resistant cell line (Panc1-GR3)
hsa-miR-4291 Gemcitabine 60750 NSC613327 approved resistant cell line (Panc1-GR4)

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